The overarching objective of MICROBES-4-CLIMATE is to provide a wider community of users/researchers, irrespective of location, with an efficient access to a cluster of complementary world-class Research Infrastructures and their integrated, advanced services, enabling them to conduct curiosity-driven, cross-disciplinary research addressing terrestrial biodiversity and ecosystems, including agricultural and (agro)forestry ecosystems, in light of the multidimensional and still poorly understood microbiomes-plants-soil-environment interactions, and its roles in climate change responses, resilience, and mitigation.